Hydrogeomorphic

Ele_COV

Coefficient of variation of elevation within a 1 km × 1 km window

DEM

Mean elevation within a 1 km × 1 km window

Dist_w

Distance to waterbodies (larger than 1 km^{2}) weighted by DEM

Dist_rv

Distance to river weighted by DEM

RvDen

River density

Human disturbance

Dist_nat

Distance to protected areas weighted by DEM

Dist_rd

Distance to roads weighted by DEM

Dist_town

Distance to settlement weighted by DEM

DamDEN

Dam density

Land productivity

COV

Coefficient of variation of monthly NDVI in a year, i.e. NDVI seasonality

NDVI.Max

Maximum NDVI in a year

NDVI.Mean

Mean monthly NDVI in a year

NDVI.Min

Minimum NDVI in a year
