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Table 1 Univariate statistics for measurements and plumage scoring of taxa of the Z. mollissimaZ. dixoni complex

From: Integrative taxonomy of the Plain-backed Thrush (Zoothera mollissima) complex (Aves, Turdidae) reveals cryptic species, including a new species

 

Alpine Thrush Z. m. whiteheadi

Alpine Thrush Z. m. mollissima s.s.

“Yunnan Thrush”

Long-tailed Thrush Z. dixoni

Sichuan Forest Thrush Z. griseiceps

Himalayan Forest Thrush Z. salimalii sp. nov.

Blyth’s type of mollissima

whiteheadi versus mollissima

All Z. mollissima s.l.  versus dixoni

Alpine versus griseiceps

Alpine versus Himalayan Forest

dixoni versus griseiceps

dixoni versus Himalayan Forest

griseiceps versus Himalayan Forest

Culmen l from skull

27.50 ± 1.22

(25.3–30.4, 21)

27.49 ± 1.00

(23.8–30.4, 93)

28.57 ± 0.23

(28.3–28.7, 3)

27.10 ± 1.24

(23.4–29.5, 33)

30.46 ± 0.59

(29.8–31.6, 8)

29.54 ± 1.44

(25.8–33.8, 53)

   

***

***

***

***

 

Culmen l from gape

29.54 ± 1.73

(26.6–34.4, 21)

29.71 ± 1.42

(24.7–33.0, 86)

31.67 ± 0.64

(31.3–32.4, 3)

29.70 ± 1.49

(26.2–32.8, 33)

33.81 ± 0.79

(32.9–35.1, 7)

31.61 ± 1.67

(27.8–36.0, 52)

28.6

  

***

***

***

***

*

Bill w from distal nares

5.25 ± 0.36

(4.6–5.7, 16)

5.47 ± 0.36

(4.7–6.2, 86)

5.77 ± 0.25

(5.5–6.0, 3)

5.34 ± 0.31

(4.6–5.9, 36)

6.12 ± 0.38

(5.5–6.6, 6)

5.96 ± 0.34

(5.2–6.8, 45)

   

***

***

***

***

 

Bill d from distal nares

5.73 ± 0.31

(5.2–5.2, 16)

5.79 ± 0.31

(4.9–6.6, 84)

6.30 ± 0.35

(5.9–6.5, 3)

5.59 ± 0.29

(5.1–6.2, 36)

7.21 ± 0.18

(7.0–7.5, 7)

6.46 ± 0.36

(5.5–7.1, 42)

   

***

***

***

***

***

Culmen ridge w

1.52 ± 0.15

(1.3–1.7, 15)

1.55 ± 0.16

(1.1–2.0, 84)

1.63 ± 0.15

(1.5–1.8, 3)

1.40 ± 0.12

(1.2–1.8, 36)

1.78 ± 0.31

(1.3–2.2, 6)

1.43 ± 0.11

(1.2–1.7, 43)

  

***

 

**

***

 

***

Bill hook l

2.36 ± 0.39

(1.7–3.2, 21)

2.29 ± 0.30

(1.5–3.0, 81)

2.37 ± 0.23

(2.1–2.5, 3)

2.02 ± 0.29

(1.3–2.6, 32)

2.60 ± 0.25

(2.2–2.9, 6)

2.58 ± 0.49

(1.3–4.2, 50)

  

**

 

**

**

***

 

Skull w

20.01 ± 0.62

(19.1–21.0, 7)

20.07 ± 0.95

(16.7–22.9, 70)

21.00 ± 0.07

(20.9–21.0, 2)

20.40 ± 1.5

(17.0–22.2, 12)

21.80 ± 0.70

(20.8–22.5, 6)

19.82 ± 0.83

(18.4–22.3, 40)

   

***

   

***

Rictal bristle l

10.07 ± 1.16

(7.1–12.7, 16)

10.68 ± 1.13

(7.7–13.8, 80)

10.63 ± 0.85

(9.8–11.5, 3)

10.81 ± 1.24

(7.5–13.6, 35)

11.62 ± 1.19

(9.9–12.9, 6)

12.33 ± 1.67

(8.9–15.4, 44)

    

***

 

***

 

Wing l flat and stretched

142.82 ± 3.46

(137.0–150.5, 25)

141.96 ± 4.46

(129.0–154.0, 94)

152.67 ± 2.52

(150.0–155.0, 3)

138.86 ± 3.77

(133.0–148.0, 38)

142.18 ± 3.72

(138.0–148.5, 8)

136.61 ± 3.80

(127.0–143.0, 51)

136.5

 

**

 

***

  

*

Wingtip l

44.18 ± 3.14

(35.0–49.0, 25)

44.73 ± 3.25

(33.0–53.0, 98)

45.33 ± 0.58

(45.0–46.0, 3)

43.19 ± 2.82

(39.0–54.0, 37)

38.83 ± 2.86

(34.0–42.0, 6)

41.24 ± 3.45

(26.0–46.0, 51)

   

***

***

   

Primary 1 (P1) shortfall

83.91 ± 2.87

(77.0–87.0, 16)

82.30 ± 4.52

(71.0–90.0, 87)

87.33 ± 0.58

(87.0–88.0, 3)

78.17 ± 2.85

(73.0–86.0, 38)

76.29 ± 3.77

(71.0–81.0, 7)

77.57 ± 4.32

(67.0–84.0, 45)

  

***

*

***

   

P2 shortfall

10.34 ± 1.98

(7.0–13.0, 16)

10.00 ± 1.85

(6.4–15.0, 86)

10.17 ± 1.04

(9.0–11.0, 3)

12.50 ± 1.85

(9.0–16.0, 38)

14.14 ± 2.56

(9.1–17.0, 8)

10.85 ± 2.096.7–15.2, 45)

9.5

 

***

***

  

*

*

P3 shortfall

0.44 ± 0.57

(0–1.5, 16)

0.44 ± 0.63

(0–2.0, 87)

0.83 ± 0.29

(0.5–1.0, 3)

1.35 ± 0.86

(0–3.3, 38)

1.75 ± 1.07

(0.5–4.0, 8)

0.91 ± 0.75

(0–2.5, 44)

0

 

***

***

**

   

P4 shortfall

0.27 ± 0.56

(0–2.0, 15)

0.16 ± 0.40

(0–2.0, 84)

0.00 ± 0.00

(3)

0.03 ± 0.16

(0–1.0, 38)

0.06 ± 0.18

(0–0.5, 8)

0.14 ± 0.41

(0–2.0, 45)

0

       

P5 shortfall

3.36 ± 1.72

(1.0–8.0, 14)

3.64 ± 1.23

(1.0–8.0, 81)

3.67 ± 0.58

(3.0–4.0, 3)

2.97 ± 0.81

(1.0–4.5, 38)

1.50 ± 0.80

(0–2.5, 8)

2.33 ± 1.13

(0–5.0, 45)

   

***

***

**

  

P6 shortfall

14.00 ± 1.7

(12.0–18.5, 15)

14.34 ± 1.72

(11.0–19.5, 81)

15.00 ± 1.00

(14.0–16.0, 3)

14.03 ± 1.5

(11.0–17.5, 36)

10.09 ± 1.67

(8.0–13.1, 7)

14.92 ± 2.04

(10.0–20.0, 45)

   

***

 

***

 

***

P7 shortfall

25.29 ± 2.20

(22.0–30.0, 14)

24.96 ± 2.09

(20.2–30.0, 80)

26.33 ± 0.58

(26.0–27.0, 3)

24.01 ± 1.44

(21.0–27.0, 36)

19.9 ± 2.57

(16.0–23.8, 7)

24.07 ± 2.44

(18.0–31.2, 45)

  

*

***

 

***

 

**

P8 shortfall

31.50 ± 3.72

(24.0–39.0, 15)

31.04 ± 2.37

(24.0–37.0, 81)

32.67 ± 0.58

(32.0–33.0, 3)

30.11 ± 1.21

(28.0–32.5, 36)

25.84 ± 2.96

(21.0–30.4, 7)

29.15 ± 2.36

(22.0–34.0, 45)

   

***

***

***

  

P9 shortfall

37.13 ± 2.77

(33.0–42.0, 15)

36.41 ± 2.84

(32.0–49.0, 81)

38.00 ± 1.00

(37.0–39.0, 3)

35.25 ± 1.27

(32.0–37.0, 36)

30.29 ± 2.14

(28.0–34.0, 7)

33.48 ± 2.15

(26.0–38.0, 44)

  

*

***

***

***

**

*

P10 shortfall

42.36 ± 3.67

(37.0–48.0, 14)

40.91 ± 2.55

(36.0–48.0, 78)

43.33 ± 1.16

(42.0–44.0, 3)

39.88 ± 1.54

(37.0–43.0, 36)

34.93 ± 1.88

(33.0–37.5, 7)

37.24 ± 2.54

(30.0–42.0, 43)

  

*

***

***

***

***

 

P3 emargination l

52.53 ± 3.20

(47.0–59.0, 16)

50.57 ± 2.90

(45.0–59.0, 81)

55.00 ± 1.00

(54.0–56.0, 3)

50.24 ± 2.49

(45.0–55.0, 37)

47.67 ± 3.08

(45.0–52.0, 6)

48.15 ± 2.71

(43.0–54.0, 43)

 

**

  

***

 

*

 

P4 emargination l

42.44 ± 2.07

(38.0–46.0, 16)

41.50 ± 2.37

(37.0–50.0, 81)

46.33 ± 2.08

(44.0–48.0, 3)

39.65 ± 2.15

(35.0–44.0, 37)

38.17 ± 1.17

(37.0–40.0, 6)

39.40 ± 2.71

(36.0–47.0, 43)

   

*

***

   

P5 emargination l

33.87 ± 1.93

(37.0–43.0, 16)

33.55 ± 2.56

(28.3–41, 80)

36.33 ± 2.52

(34.0–39.0, 3)

31.03 ± 3.03

(25.0–39.0, 36)

31.00 ± 2.19

(29.0–35.0, 6)

31.18 ± 2.40

(24.0–37.0, 43)

    

***

   

P2 notch l

38.88 ± 1.93

(37.0–43.0, 16)

38.99 ± 2.86

(29.0–51.0, 78)

40.00 ± 2.65

(37.0–42.0, 3)

35.97 ± 2.73

(26.0–41.0, 37)

35.00 ± 2.19

(32.0–38.0, 6)

35.40 ± 2.60

(30.0–42.0, 37)

  

**

 

***

   

P3 notch l

37.73 ± 2.22

(34.0–4.01, 15)

36.77 ± 2.51

(29.0–42.0, 75)

41.00 ± 1.00

(40.0–42.0, 3)

34.13 ± 2.08

(30.0–38.0, 32)

33.67 ± 1.51

(32.0–35.0, 6)

33.71 ± 2.16

(29.0–38.0, 35)

  

***

 

***

   

Tail l

103.94 ± 4.90

(92.0–113.0, 24)

103.25 ± 4.54

(92.0–114.0, 99)

114.0 ± 1.73

(113.0–116.0, 3)

114.61 ± 5.20

(104.0–126.0, 36)

111.75 ± 4.03

(106.0–116.0, 8)

94.49 ± 4.93

(81.7–106.0, 51)

101.6

 

***

***

***

 

***

***

Undertail coverts to tail tip

35.76 ± 3.14

(31.0–42.0, 14)

36.19 ± 5.16

(28.0–58.0, 80)

37.00 ± 2.65

(35.0–40.0, 3)

45.13 ± 4.68

(33.0–58.2, 36)

43.67 ± 6.09

(40.0–56.0, 6)

34.33 ± 5.07

(25.8–47.0, 43)

  

***

*

  

***

**

Uppertail coverts to tail tip

42.03 ± 7.63

(28.0–56.0, 13)

40.73 ± 5.66

(31.0–60.0, 82)

41.00 ± 2.83

(39.0–43.0, 2)

57.97 ± 5.33

(51.0–71.5, 30)

52.33 ± 4.55

(49.0–61.0, 6)

43.75 ± 4.68

(35.0–54.0, 42)

  

***

***

  

***

**

Tail graduation

2.37 ± 2.10

(0–7.0, 15)

2.71 ± 1.62

(0–7.0, 80)

2.33 ± 1.53

(1.0–4.0, 3)

5.00 ± 2.02

(1.8–11.0, 32)

5.17 ± 0.75

(4.0–6.0, 6)

3.43 ± 1.52

(1.0–6.1, 40)

  

***

*

  

*

 

Maximum w of inner rectrix

17.69 ± 1.61

(15.0–20.0, 13)

17.10 ± 1.41

(13.0–22.0, 79)

17.3 ± 0.58

(17.0–18.0, 3)

17.73 ± 1.29

(15.0–21.0,33)

19.33 ± 1.51

(17.0–21.0, 6)

17.41 ± 1.26

(15.0–20.0, 40)

   

*

    

Maximum l of white on outer rectrix

11.54 ± 4.71

(3.0–21.0, 17)

12.54 ± 5.88

(2.0–33.0, 81)

15.00 ± 5.00

(10.0–20.0.5, 3)

25.49 ± 6.95

(16.0–52.0, 33)

12.33 ± 3.50

(8.0–18.0, 6)

14.81 ± 4.53

(8.0–28.0, 43)

  

***

  

**

***

 

Minimum l of white on outer rectrix

3.28 ± 0.87

(2.0–6.0, 17)

3.55 ± 1.03

(1.0–7.0, 81)

3.00 ± 1.00

(2.0–4.0, 3)

12.00 ± 3.76

(4.0–20.0, 32)

3.83 ± 2.56

(1.0–7.0, 6)

4.14 ± 2.29

(2.0–13.0, 41)

  

***

  

**

***

 

Tarsus l

37.16 ± 1.10

(35.4–39.1, 25)

36.91 ± 1.20

(33.8–39.9, 99)

37.67 ± 1.23

(36.3–38.7, 3)

37.70 ± 0.90

(35.6–39.7, 38)

41.31 ± 1.60

(39.3–43.5, 8)

34.00 ± 1.37

(31.2–38.1, 53)

34.9

  

***

***

***

***

***

Hindclaw l

10.33 ± 0.64

(8.8–11.2, 19)

10.30 ± 0.51

(9.2–11.3, 84)

10.6 ± 0.00

(3)

9.56 ± 0.53

(8.4–10.6, 38)

10.51 ± 0.53

(9.7–11.2, 8)

10.19 ± 0.59

(8.9–11.6, 46)

  

***

  

**

***

 

Bill l/wing l ratio

0.19 ± 0.01

(0.2–0.2, 22)

0.19 ± 0.01

(0.1–0.2, 80)

0.19 ± 0.01

(0.2–0.2, 3)

0.20 ± 0.01

(0.2–0.2, 32)

0.21 ± 0.01

(0.2–0.2, 8)

0.22 ± 0.01

(0.2–0.2, 48)

   

***

***

***

***

 

Wing l/tail l ratio

1.38 ± 0.05

(1.3–1.5, 24)

1.38 ± 0.05

(1.3–1.6, 92)

1.34 ± 0.04

(1.3–1.4, 3)

1.22 ± 0.05

(1.1–1.3, 37)

1.27 ± 0.03

(1.2–1.3, 8)

1.44 ± 0.07

(1.3–1.6, 46)

  

***

***

***

 

***

***

Bill l/tarsus l ratio

0.74 ± 0.04

(0.7–0.8, 22)

0.74 ± 0.03

(0.7–0.8, 87)

0.76 ± 0.03

(0.7–0.8, 3)

0.72 ± 0.03

(0.6–0.8, 32)

0.74 ± 0.04

(0.7–0.8, 8)

0.87 ± 0.06

(0.7–1.0, 50)

  

*

 

***

 

***

***

Bill l/culmen ridge w ratio

18.09 ± 1.75

(15.9–21.5, 13)

17.56 ± 1.47

(14.4–19.8, 58)

 

19.37 ± 1.84

(15.1–24.6, 31)

18.36 ± 2.86

(15.9–23.1, 5)

21.07 ± 1.96

(16.2–26.0, 37)

  

**

 

***

 

**

 

Bill l/bill d ratio

4.78 ± 0.29

(4.4–5.5, 13)

4.74 ± 0.22

(4.2–5.3, 67)

4.54 ± 0.22

(4.4–4.8, 3)

4.86 ± 0.30

(4.2–5.44, 31)

4.23 ± 0.10

(4.0–4.3, 7)

4.59 ± 0.21

(4.1–4.9, 38)

   

***

*

***

**

**

Tarsus l/hallux l ratio

3.62 ± 0.23

(3.3–4.0, 19)

3.60 ± 0.18

(3.3–4.1, 53)

3.55 ± 0.12

(3.4–3.6, 3)

3.95 ± 0.21

(3.6–4.5, 38)

3.95 ± 0.22

(3.7–4.3, 7)

3.32 ± 0.23

(2.8–3.9, 41)

  

***

**

***

 

***

***

Wing l/tarsus l ratio

3.85 ± 0.14

(3.6–4.2, 25)

3.84 ± 0.13

(3.6–4.1, 91)

4.05 ± 0.09

(4.0–4.1, 3)

3.69 ± 0.12

(3.5–3.9, 38)

3.44 ± 0.09

(3.3–3.6, 8)

4.03 ± 0.20

(3.7–4.4, 48)

  

***

***

***

***

***

***

Mandible color (1 = all fleshy–10 = all black)

5.77 ± 1.69

(3–8, 13)

6.85 ± 1.15

(3–9, 61)

8.67 ± 0.58

(8–9, 3)

6.10 ± 0.87

(4–8, 31)

7.25 ± 1.03

(5–8, 8)

8.14 ± 1.10

(5–10, 42)

 

*

*

 

***

**

***

*

Crown color (1 = medium brown–10 = very dark brown)

2.71 ± 1.21

(1–5, 17)

5.10 ± 1.58

(2–8, 60)

8.00 ± 1.73

(6–9, 3)

5.50 ± 1.01

(3–7, 30)

8.13 ± 1.13

(6–9, 8)

7.48 ± 0.97

(5–9, 42)

 

***

 

***

***

***

***

 

Crown value (1 = grey-brown–10 = russet)

3.06 ± 1.30

(1–5, 17)

4.02 ± 1.37

(2–7, 60)

2.33 ± 1.16

(1–3, 3)

5.10 ± 0.96

(3–7, 30)

2.00 ± 0.76

(1–3, 8)

7.44 ± 1.14

(4–9, 41)

 

**

***

***

***

***

***

***

Crown scalloping (1 = uniform–10 = dark scalloping)

2.41 ± 0.94

(1–4, 17)

4.17 ± 1.44

(1–7, 59)

5.67 ± 1.53

(4–7, 3)

2.50 ± 0.97

(1–5, 30)

6.25 ± 1.49

(4–8, 8)

3.22 ± 2.19

(1–7, 41)

 

*

***

***

*

***

 

**

Crown-mantle contrast (1 = none–10 = strong)

2.06 ± 0.90

(1–3, 17)

3.06 ± 1.08

(2–8, 32)

7.33 ± 0.58

(7–8, 3)

2.60 ± 0.72

(1–4, 30)

7.00 ± 1.53

(4–8, 7)

2.20 ± 0.52

(1–3, 20)

    

***

***

 

***

Mantle color (1 = very olive–10 = very russet)

3.06 ± 1.30

(1–5, 17)

4.21 ± 1.47

(2–6, 33)

6.33 ± 1.53

(5–8, 3)

4.07 ± 0.83

(3–6, 30)

6.83 ± 1.17

(6–9, 6)

7.95 ± 0.59

(7–9, 21)

 

*

 

**

***

***

***

**

Eyering color (1 = white–5 = buff–10 = absent)

2.00 ± 1.00

(1–5, 17)

2.71 ± 0.99

(1–5, 48)

3.67 ± 1.16

(3–5, 3)

1.75 ± 0.70

(1–3, 28)

3.29 ± 1.25

(2–5, 7)

4.12 ± 0.87

(2–5, 41)

  

***

 

***

**

***

 

Eyering breadth (1 = very broad–10 = absent)

2.94 ± 1.48

(1–5, 17)

3.02 ± 1.07

(1–5, 49)

3.50 ± 2.12

(2–5, 2)

2.93 ± 1.33

(1–6, 28)

5.00 ± 1.29

(3–7, 7)

4.46 ± 1.14

(2–7, 41)

   

***

***

**

***

 

Pale supraloral extent (1 = much pale–10 = no pale)

3.77 ± 1.79

(1–8, 17)

3.22 ± 1.50

(1–8, 59)

3.00 ± 1.00

(2–4, 3)

2.23 ± 0.85

(1–4, 31)

3.25 ± 2.05

(2–8, 8)

5.55 ± 1.85

(2–9, 42)

  

**

 

***

 

***

**

Lores value (1 = all-pale–10 = all dark)

5.29 ± 1.49

(3–8, 17)

4.72 ± 1.46

(2–10, 57)

6.33 ± 1.16

(5–7, 3)

3.77 ± 0.97

(2–5, 30)

5.86 ± 1.21

(4–7, 7)

8.17 ± 1.27

(4–10, 42)

  

**

 

***

***

***

***

Auriculars blotchiness (1 = uniform–10 = large blotches)

4.41 ± 1.87

(1–7, 17)

5.71 ± 1.08

(3–8, 58)

4.50 ± 3.54

(2–7, 2)

8.00 ± 1.07

(6–10, 31)

2.00 ± 0.82

(1–3, 7)

4.98 ± 1.77

(1–8, 41)

  

***

***

**

***

***

***

Auriculars streakiness (1 = streaked–10 = unstreaked)

4.47 ± 1.7

(1–7, 17)

4.43 ± 1.58

(1–8, 58)

4.50 ± 3.54

(2–7, 2)

5.73 ± 2.0

(3–9, 30)

1.57 ± 0.79

(1–3, 7)

3.46 ± 1.34

(2–7, 41)

   

***

***

***

***

***

Pale submoustachial prominence (1 = not apparent–10 = all-white)

4.00 ± 2.24

(2–10, 17)

4.90 ± 1.54

(2–10, 58)

5.33 ± 0.58

(5–6, 3)

3.74 ± 1.29

(2–7, 31)

7.12 ± 0.99

(6–9, 8)

6.17 ± 1.21

(3–8, 42)

 

*

*

***

***

***

***

 

Dark malar prominence (1 = not apparent, 10 = black)

4.06 ± 1.35

(3–7, 17)

5.19 ± 1.33

(3–9, 58)

4.00 ± 1.73

(2–5, 3)

3.97 ± 1.05

(2–6, 31)

6.38 ± 2.07

(4–9, 8)

7.45 ± 1.31

(5–10, 42)

 

*

**

 

***

**

***

 

Throat color (1 = white–10 = buff)

2.59 ± 1.46

(1–6, 17)

3.75 ± 1.60

(1–8, 59)

4.33 ± 0.58

(4–5, 3)

2.19 ± 1.05

(1–7, 31)

5.88 ± 1.55

(3–8, 8)

6.50 ± 1.50

(3–9, 42)

 

**

***

***

***

***

***

 

Throat marking (1 = none–10 = heavy)

3.41 ± 1.50

(2–7, 17)

4.09 ± 1.47

(2–8, 59)

6.00 ± 2.65

(3–8, 3)

2.58 ± 0.85

(1–6, 31)

2.25 ± 0.89

(1–4, 8)

2.67 ± 0.95

(1–6, 42)

 

*

***

***

***

   

Breast ground color (1 = white–10 = buff)

5.77 ± 2.56

(2–9, 17)

6.70 ± 1.60

(2–10, 60)

8.33 ± 1.16

(7–9, 3)

4.23 ± 1.33

(2–9, 31)

8.88 ± 0.64

(8–10, 8)

8.00 ± 0.86

(6–9, 42)

 

*

***

***

***

***

***

*

Central breast mark shape (1 = triangles–10 = chevrons)

1.88 ± 1.17

(1–5, 17)

1.49 ± 1.15

(1–8, 59)

4.33 ± 0.58

(4–5, 3)

3.61 ± 1.02

(1–5, 31)

3.25 ± 1.49

(1–5, 8)

2.57 ± 1.21

(1–7, 42)

  

**

***

***

 

***

 

Central breast mark density (1 = very light–10 = very heavy)

5.35 ± 2.23

(1–8, 17)

6.08 ± 1.78

(1–9, 60)

6.00 ± 1.00

(5–7, 3)

4.55 ± 1.15

(2–7, 31)

5.00 ± 1.51

(3–7, 8)

7.21 ± 1.24

(4–9, 42)

  

**

 

***

 

***

***

Flank mark density (1 = very light–10 = very heavy)

4.29 ± 1.40

(2–7, 17)

5.12 ± 1.60

(2–8, 60)

7.33 ± 0.58

(7–8, 3)

4.03 ± 1.74

(2–8, 31)

5.38 ± 2.00

(2–8, 8)

5.59 ± 1.48

(3–8, 42)

 

*

**

   

***

 

Flank mark shape (1 = straight–10 = chevrons)

6.88 ± 2.18

(3–10, 17)

7.07 ± 1.46

(2–9, 60)

5.68 ± 1.53

(4–7, 3)

1.36 ± 0.55

(1–3, 31)

4.00 ± 2.27

(2–7, 8)

4.79 ± 1.98

(1–10, 42)

 

**

***

***

***

***

***

 

Central belly markings (1 = none–10 = heavy)

3.41 ± 1.23

(2–7, 17)

3.27 ± 1.00

(2–6, 56)

2.50 ± 0.71

(2–3, 2)

1.29 ± 0.46

(1–2, 31)

2.50 ± 0.76

(2–4, 8)

3.83 ± 1.22

(3–7, 42)

  

***

*

*

***

***

***

Vent percent dark (1 = all-white 10 = all dark)

5.41 ± 1.18

(4–7, 17)

4.66 ± 1.25

(1–8, 61)

4.67 ± 0.58

(4–5, 3)

2.45 ± 0.51

(2–3, 31)

6.25 ± 1.16

(5–8, 8)

6.52 ± 1.31

(5–9, 42)

 

*

***

**

***

***

***

 

Undertail-tail tips contrast (1 = very weak–10 = very strong)

3.19 ± 0.91

(2–5, 16)

3.11 ± 1.31

(2–8, 65)

6.33 ± 2.89

(3–8, 3)

4.55 ± 1.45

(2–7, 29)

6.00 ± 1.77

(3–8, 8)

7.24 ± 1.38

(3–9, 42)

  

**

***

***

*

***

*

Claw color (1 = much darker than toes–10 = much paler than toes)

3.69 ± 1.12

(2–5, 17)

3.37 ± 1.13

(1–7, 59)

2.67 ± 1.53

(2–5, 3)

6.23 ± 1.23

(3–8, 31)

7.88 ± 1.13

(7–10, 8)

7.40 ± 1.35

(3–10, 42)

  

***

***

***

**

***

 

Patterning on folded wing (1 = none–10 = strong)

3.88 ± 1.27

(2–5, 17)

6.12 ± 1.02

(4–8, 33)

3.33 ± 1.53

(2–5, 3)

7.23 ± 0.96

(5–9, 31)

3.29 ± 1.35

(2–6, 7)

4.57 ± 0.93

(2–6, 41)

  

***

***

***

***

***

**

Primary coverts pattern (1 = none–10 = strong)

5.00 ± 1.37

(3–7, 17)

6.97 ± 0.98

(4–8, 33)

4.33 ± 2.52

(2–7, 3)

7.68 ± 0.70

(6–9, 31)

4.00 ± 1.58

(2–6, 5)

5.81 ± 1.08

(4–8, 21)

  

***

***

***

***

***

**

Secondary covert edge width (1 = broad–10 = absent)

7.12 ± 1.27

(4–9, 17)

7.51 ± 1.46

(4–10, 33)

7.33 ± 1.16

(6–8, 3)

2.61 ± 1.05

(1–7, 31)

4.71 ± 1.50

(3–7, 7)

8.38 ± 1.16

(5–10, 21)

 

*

***

***

**

***

***

***

Secondary covert edge color (1 = white–5 = buff–10 = absent)

5.77 ± 2.25

(2–9, 17)

5.52 ± 2.59

(1–10, 33)

5.33 ± 2.08

(3–7, 3)

4.87 ± 0.34

(4–5, 31)

5.71 ± 1.25

(5–8, 7)

6.09 ± 1.81

(4–10, 21)

     

*

***

 

Tertial edge strength (1 = strong–10 = absent)

7.27 ± 2.05

(3–10, 15)

8.36 ± 1.08

(4–9, 33)

6.00 ± 2.65

(3–8, 3)

6.35 ± 2.08

(3–10, 29)

8.29 ± 0.76

(7–9, 7)

8.71 ± 1.49

(3–10, 21)

    

*

*

***

 

Tertial edge color (1 = white–5 = buff–10 = absent)

5.53 ± 3.07

(1–10, 15)

6.00 ± 3.04

(1–9, 33)

4.33 ± 3.22

(2–8, 3)

6.00 ± 2.2

(3–10, 29)

8.00 ± 1.87

(5–10, 7)

8.38 ± 1.75

(3–10, 21)

  

*

 

***

 

***

 
  1. Significance levels (* P < 0.05; ** P < 0.01; *** P < 0.001; all others not significant)
  2. l length, w width, d depth